After the “reboot” and Meicoomon’s rampage, Tai and friends arrive in the Digital World. They reunite with their partner Digimon, who have lost all their memories. As everyone discusses what they should do from here in the Digital World, Meicoomon suddenly appears and then disappears. She still has her memories for some reason as she wanders around, looking for Meiko.

Meanwhile in the real world, Nishijima receives word that Himekawa has gone missing. As he investigates, he determines that there’s been some hidden agenda behind her behavior up to this point. It has something to do with an event in the past that determined both their destinies… The adventure evolves once again.